A review of current knowledge of the distribution of Palaeozoic ferns and fern-like plants is presented. Whilst numerous putative ferns occur in the Devonian, it is not until the Lower Carboniferous that ferns belonging to the Filicales appear. The Namurian represents a major gap in our knowledge. The ecology of late Palaeozoic ferns is reviewed. The association of early fern assemblages with volcanic terrains is considered particularly significant in the relationship between evolution and environment. Extinction of more specialised groups may have been caused by climatic changes.